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Jim, gave good info!!
But to break it down as far as dates a little clearer for you I thought I'd share a little more insight to help you with your hunting decisions.
There are 3 phases of the Rut, many years a 4th. The 1st Aug. 20th to Sept.10th, this is when the first cows usually will show signs, it's the older cows only that do, therefore it can be sporatic at best as this is not peak rut, nonetheless bulls can get aggressive at this time, a lot to a little bugling is possible if you're in the right spot/area.
The 2nd phase is "peakrut" this happens from the 17th-25th of Sept. This is when Most Cows come into estrus, it's also peak bugling time! Cows are in heat for 6-12 hours.
The 3rd phase is generally around the 15th of Oct. & occurs everyear, this rut takes part for cows that haven't been covered or didn't take the first time around. It's not nearly as intense as peak rut, but still a great time to be in the woods. These times of estrus are generally 21 days apart.
A 4th can take place in early Nov. but is a crap shoot & not to be relied on. Once again for cows that haven't took! This is why you'll hear some years where bulls were going bonkers over calls in selected areas not everywhere.
You can expect bugling during & between the first 3 phases. THe further from the roads/trails you are after peak rut & hunting pressure the better your chances of catching responsive to calls elk.
Hope this helps out to pick the right time for what you expect out of your hunt. Good Luck!
ElkNut1